Plants: moderately large, densely caespitose, golden yellow to golden brown, somewhat glossy. Stems: creeping, densely or rarely loosely pinnate; hyalodermis absent, central strand present, weak; pseudoparaphyllia foliose, lanceolate, never filamentous. Stem: and branch leaves with dorsal, lateral, and ventral stem leaves often different in shape and size. Stem: leaves closely imbricate, narrowly lanceolate, plicate; base not or short- to long-decurrent; margins recurved at base, entire to subentire proximally, serrate, serrulate, or rarely entire distally; apex narrowly acute [blunt]; costa double; alar cells differentiated, subquadrate to rectangular; laminal cells smooth, walls thick; medial cells linear. Branch: leaves much smaller, narrower. Specialized: asexual reproduction absent. Sexual: condition dioicous; inner perichaetial leaves oblong-lanceolate, apex filiform-acuminate. [Seta reddish brown. Capsule: inclined to horizontal, ovoid to cylindric, straight or weakly curved, not or weakly contracted below mouth; annulus little differentiated; operculum conic, sometimes short-rostrate; peristome double; exostome teeth with external surface transversely smooth to striolate proximally, papillose distally; endostome basal membrane 1/2 exostome teeth length, segments keeled, cilia 2â4. Calyptra: naked. Spores: rounded, finely papillose]. Distribution: nw North America, Asia (Bhutan, China, India, Indonesia, Japan, Korea, Nepal, Pakistan, e Russia, Taiwan), Pacific Islands (Philippines).
Discussion: Species 20 (1 in the flora).